This feature only applies when the pileus trama is dimitic, with generative hyphae and a second type of strengthening hyphae. These secondary hyphae are usually unbranched (skeletal hyphae) but may be branched (skeleto-ligative and binding hyphae).

Choose this state if: the secondary hyphae in the pileus trama are in the form of skeleto-ligative or binding hyphae, both of which are branched, sometimes highly so.

Skeleto-ligative hyphae, which among lamellate fungi are particularly characteristic of the genus Lentinus, are also called skeleto-binding cells. Skeleto-ligative hyphae are thick-walled and don't have any cross walls. They have fine, tapering side branches, which may be further branched. There are usually several main branches, one of which is directed backwards at the base and another forwards near the apex of the main axis. The side branches are referred to as binding processes, or binding or ligative hyphae (although the branches are in fact not separate hyphae but are continuous with the main axis of the hypha).

Binding hyphae are highly branched, sometimes with an unbranched basal portion (when they are referred to as arboriform skeletal hyphae).