This feature only applies when the pileus trama is dimitic, with generative hyphae and a second type of strengthening hyphae. These secondary hyphae are usually unbranched (skeletal hyphae) but may be branched (skeleto-ligative and binding hyphae).

Choose this state if: the secondary hyphae in the pileus trama are in the form of skeletal hyphae. These are thick-walled and aseptate without clamp connections, although they may be clamped at the base. They are unbranched, or if branches are present, they are few and do not taper.