The direction of the lamellar trama refers to the overall direction of hyphae within the lamellar trama, whether at right angles (descending) or parallel (radiate) to the edge of the lamella. Assess this feature from the same section used to ascertain Lamellar trama (pattern), which is a cross section of a lamella running from the edge to the top, at right angles to the edge. This feature is best observed on young material, since in older fruit-bodies a descending arrangement can become irregular, although it is usually retained at the edge of the lamella.

Choose this state if: the hyphae of the lamellar trama are radially arranged, more or less parallel to the edge of the lamellae. A cross section of the lamellar trama at right angles to the lamellae edge will show most hyphae in cross section (circular).

Where the lamellar trama is radiate, the base of the lamella often readily separates from the pileus, and the edge of the lamella also is readily separable.